• forewing vein R5 free
• forewing with oval sclerotization costad of base of vein A1+2
• bullae tympani open cephalad
• tympanum and conjunctivum lying at a blunt angle
• processus tympani present
• praecinctorium present
• accessory tympana present caudally of metacoxae
• male genitalia without uncus arms
• segment A8 of larvae without sclerotized ring around base of SD1